-ulo

Etymology
Probably from and, both feminine noun suffixes.

Suffix

 * 1)  member of group of animals characterized by the root
 * 1)  member of group of animals characterized by the root
 * 1)  member of group of animals characterized by the root
 * 1)  member of group of animals characterized by the root
 * 1)  member of group of animals characterized by the root

Etymology
.